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Course Catalog 2003-2004

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Division of Biological Sciences

Donald P. Christian, Associate Dean for the Biological Sciences
The Division of Biological Sciences offers undergraduate and graduate programs representing the full range of the biological sciences. The Division offers bachelor degrees in biology (with a broad array of formal options including botanical sciences, cellular and molecular biology, ecology, human biological sciences, natural history, zoological sciences, biological education and general sciences education), medical technology, microbiology including microbial ecology, and wildlife biology (a cooperative program administered by the College of Forestry and Conservation). The Division also advises students in pre-medical and other pre-health sciences, and offers a series of courses during the summer at the University's Flathead Lake Biological Station (see separate listing in this section). The Division is one of the leading research units in the University. Research programs in the Division provide abundant opportunities for students to enhance their educational experience by participating in research. Several sources of funding are available to support undergraduate student research, and the Division participates in the University undergraduate research symposium each spring.

Graduate degrees offered by the Division of Biological Sciences include Master of Science degrees in biochemistry, microbiology, and organismal biology and ecology. Doctor of Philosophy degrees in biochemistry/microbiology, organismal biology and ecology, and biomolecular structure and dynamics are offered. The Division participates in the graduate (M.S. and Ph.D.) program in wildlife biology, administered by the College of Forestry and Conservation. Information on graduate study and program requirements is available from the Graduate School or the Division of Biological Sciences.

The Division of Biological Sciences is committed to providing coursework and experiences for non-science majors. The world faces many problems and opportunities that include significant biological components. Courses for non-science majors have the goal of fostering understanding of the process of science and enhancing biological knowledge as it relates to environmental, medical, social, and other issues. A number of introductory courses are open both to majors and non-majors. In addition, the Division offers courses designed specifically for non-majors: elementary medical microbiology, introductory ecology, Montana wildlife, and others.

Degree requirements and courses are described below (see the College of Forestry and Conservation for information about wildlife biology).



Fred W. Allendorf, Ph.D., University of Washington, 1975
Ragan M. Callaway, Ph.D. University of California at Santa Barbara, 1990
Donald P. Christian, Ph.D., Michigan State University, 1977 (Associate Dean)
Kenneth P. Dial, Ph.D., Northern Arizona University, 1984
Kerry R. Foresman, Ph. D., Uni­versity of Idaho, 1977
James E. Gannon, Ph.D., University of Houston, 1981
Willard O. Granath, Ph.D., Wake Forest University, 1982
William E. Holben, Ph.D., University of New York, Buffalo, 1985
Richard L. Hutto, Ph.D., University of California at Los Angeles, 1977
Ralph C. Judd, Ph.D., The University of Montana, 1979
Michael F. Minnick, Ph.D., Washington State University, 1987
Jack H. Nunberg, Ph.D., Stanford University, 1979 (Director for the Montana Biotechnology Center)
Jack A. Stanford, Ph.D., University of Utah, 1975 (Bierman Professor; Director of the Biological Station)

Associate Professors

Carol A. Brewer, Ph.D., University of Wyoming, 1993
Douglas Emlen, Ph.D., Princeton University, 1994
Erick P. Greene, Ph.D., Princeton Uni­versity, 1989
Mark L. Grimes, Ph.D., University of Oregon, 1986
J. Stephen Lodmell, Ph.D., Brown University, 1996
John L. Maron, Ph.D., University of California-Davis, 1996
Mary L. Poss, Ph.D., Colorado State University, 1990
Matthias Rillig, Ph.D., San Diego State University, 1997
Frank Rosenzweig, Ph.D., University of Pennsylvania, 1991
Anna M. Sala, Ph.D., University of Barcelona, 1992
D. Scott Samuels, Ph.D., University of Arizona, 1991

Assistant Professors

Lila Fishman, Ph.D., Princeton University, 1998
Christopher G. Guglielmo, Ph.D., Simon Fraser University, 1999
Michele A. McGuirl, Ph.D., Montana State University, 1999
Scott R. Miller, Ph.D., University of Oregon, 1999


Kathleen A. Westphal, Ph.D., University of Kentucky, 1985

Research Professors

Jerry J. Bromenshenk, Ph.D., Montana State University, 1973
F. Richard Hauer, Ph.D., North Texas State University, 1980
Penelope F. Kukuk, Ph.D., University of Kansas, 1980
Barbara E. Wright, Ph.D., Stan­ford University, 1951

Research Associate Professors

Roland L. Redmond, Ph.D., The University of Montana, 1984

Research Assistant Professors

Vijay Gadkar, Ph.D., University of Delhi, 1999
Matthew J. Kauffman, Ph.D., University of California at Santa Cruz, 2003
John Kimball, Ph.D., Oregon State University, 1995
Mark Lorang, Ph.D., Oregon State University 1997
Yolanda E. Morbey, Ph.D., Simon Fraser University, 2001­
Paul Spruell, Ph.D., Washington State University, 1994

Other Adjunct Faculty

I. Joe Ball, Ph.D., University of Minnesota, 1973
Richard J. Bridges, Ph.D., Cornell University Graduate School of Medical Sciences, 1984
Barry N. Brown, M.S., University of Alaska, Fairbanks, 1987
David T. Cheung, Ph.D., University of Southern California, 1981
Elizabeth Crone, Ph.D., Duke University, 1995
Vernon R. Grund, Ph.D., University of Minnesota Medical School, 1974
Craig A. Johnston, Ph.D., Michigan State University, 1982
Charles T. Leonard, Ph.D., Medical College of Pennsylvania, 1985
Thomas E. Martin, Ph.D., University of Illinois, 1982
George Stanley, Ph.D., University of Kansas, 1977

Emeritus Professors

Mark J. Behan, Ph.D., University of Washington, 1963
David E. Bilderback, Ph.D., University of Oregon, 1968
George L. Card, Ph.D., University of Nebraska, 1966
H. Richard Fevold, Ph.D., University of Utah, 1961
James R. Habeck, Ph.D., University of Wisconsin, 1959
Walter E. Hill, Ph.D., University of Wisconsin, 1967
Donald A. Jenni, Ph.D., University of Florida, 1961
Delbert L. Kilgore, Ph.D., University of Kansas, 1972
Galen P. Mell, Ph.D., University of Washington, 1961
Lee H. Metzgar, Ph.D., University of Michigan, 1968
Charles N. Miller, Ph.D., University of Michigan, 1965
Andrew L. Sheldon, Ph.D., Cornell University, 1966
Richard P. Sheridan, Ph.D., University of Oregon, 1966
John F. Tibbs, Ph.D., University of Southern California, 1968



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